:root{color-scheme:dark;--bg:#10120f;--surface:#191d16;--line:#363e31;--text:#f3f5ed;--muted:#bec5b6;--lime:#c5f668;--amber:#ffcf87}
*,*::before,*::after{box-sizing:border-box}
[hidden]{display:none!important}
body{margin:0;background:var(--bg);color:var(--text);font:1rem/1.6 system-ui,sans-serif}
a{color:var(--lime);text-underline-offset:.2em}
a:focus-visible,input:focus-visible,select:focus-visible{outline:3px solid var(--lime);outline-offset:4px}
img{max-width:100%;display:block}
p,figure{margin:0 0 .75rem}
h1,h2,h3,h4{margin:0 0 .6rem;line-height:1.12;letter-spacing:-.025em}
h1{font-size:clamp(2.7rem,7vw,6rem);letter-spacing:-.066em}
h1 em{color:var(--lime);font-style:normal}
h2{font-size:clamp(1.45rem,3.5vw,2.35rem)}
h3,h4{font-size:1.25rem}
.wrap{width:min(1100px,calc(100% - 2rem));margin-inline:auto}
.bar{display:flex;align-items:center;justify-content:space-between;gap:.6rem 1.5rem;flex-wrap:wrap;min-height:70px}
.mast,.foot{background:#0d0f0c;border-bottom:1px solid var(--line)}
.mast nav{display:flex;gap:.35rem 1.3rem;flex-wrap:wrap;font-size:.89rem}
.mast nav a{color:var(--text);font-weight:650;text-decoration:none}
.mast nav a:hover,.mast nav [aria-current]{color:var(--lime);text-decoration:underline}
.brand{display:inline-flex;align-items:center;gap:.55rem;color:var(--text);font-weight:850;letter-spacing:.04em;text-decoration:none}
.brand b{color:var(--lime)}
.skip{position:absolute;z-index:2;inset:.5rem auto auto 1rem;padding:.6rem;background:var(--bg);transform:translateY(-200%)}
.skip:focus{transform:none}
.preview{margin:1rem 0 0;color:var(--amber);font-size:.81rem}
.hero{display:grid;grid-template-columns:minmax(0,1fr) auto;align-items:center;gap:2rem;padding:clamp(2rem,6vw,5rem) 0}
.hero.split{grid-template-columns:minmax(0,1.15fr) minmax(0,.85fr)}
.hero.detail{padding-top:1.75rem;padding-bottom:2rem}
.hero figure{margin:0}
.hero img{width:100%;height:auto;aspect-ratio:8/5;object-fit:cover;border:1px solid var(--line);border-radius:.75rem;background:var(--surface)}
.hero-index{display:grid;gap:.4rem;background:radial-gradient(ellipse at right,#26371d,var(--surface) 75%);padding:1.45rem 1.7rem;border:1px solid var(--line);border-radius:.65rem;min-width:13.5rem}
.hero-index span,.eyebrow{color:var(--lime);font-size:.76rem;font-weight:800;letter-spacing:.13em;text-transform:uppercase}
.hero-index strong{font-size:4.8rem;color:var(--text);line-height:1;letter-spacing:-.09em}
.lede{font-size:clamp(1.05rem,2vw,1.25rem);max-width:54ch;color:var(--muted)}
.latin{font-style:italic;color:var(--lime)}
.crumbs{font-size:.88rem;color:var(--muted);padding-top:1.15rem}
.crumbs span{padding-inline:.25rem}
.filters{display:flex;flex-wrap:wrap;gap:.75rem 1rem;align-items:end;margin:1rem 0 .35rem}
.filters label{display:grid;gap:.2rem;font-size:.87rem;font-weight:650}
.filters label:first-child{flex:1 1 17rem;max-width:30rem}
.filters>div{display:grid;gap:.2rem}
.filters input,.filters select{font:inherit;color:var(--text);background:var(--surface);border:1px solid var(--line);padding:.55rem .7rem;border-radius:.4rem;min-width:0;min-height:44px}
.filters input::placeholder{color:var(--muted)}
.count{color:var(--muted);font-size:.88rem}
.count:empty{display:none}
.cards,.rows,.references{padding:0;margin:0;list-style:none}
.cards{display:grid;grid-template-columns:repeat(auto-fit,minmax(min(100%,17rem),1fr));gap:1rem}
.morph-cards{grid-template-columns:repeat(auto-fit,minmax(min(100%,15rem),1fr))}
.cards:not(.morph-cards){grid-template-columns:repeat(auto-fill,minmax(min(100%,22rem),1fr))}
.species-card{display:flex;flex-direction:column;border:1px solid var(--line);border-radius:.75rem;background:var(--surface);overflow:hidden}
.species-card:hover,.species-card:focus-within{border-color:var(--lime)}
.species-card a:first-child{display:block;color:inherit;text-decoration:none;flex:1}
.species-card a:first-child:focus-visible{outline-offset:-4px}
.species-card a:first-child:hover :is(h3,h4){text-decoration:underline;text-underline-offset:.2em}
.species-card img,.photo-empty{width:100%;height:auto;aspect-ratio:8/5;object-fit:cover;background:#232b1f}
.species-card:not(.morph-card) img{aspect-ratio:13/6}
.species-card img.species-card-illustration,.hero img.species-card-illustration{object-fit:contain;background:#fff}
.photo-empty{position:relative;display:grid;place-items:center;padding-bottom:1.5rem;color:var(--muted);font-size:.8rem;background:radial-gradient(ellipse at 50% 42%,#2a3127,#1d231b 75%)}
.photo-empty>img{width:72%;height:auto;aspect-ratio:auto;object-fit:contain;border:0;border-radius:0;background:none}
.photo-empty>span{position:absolute;inset:auto 0 .65rem;text-align:center}
.hero .photo-empty{border:1px solid var(--line);border-radius:.75rem}
.card-body{padding:1rem 1.1rem .45rem}
.card-body :is(h3,h4) span,.arrow{color:var(--lime);font-size:.85em;margin-left:.35em}
.card-body p:not(.eyebrow),.card-body i{color:var(--muted)}
.credit,figcaption{font-size:.77rem;color:var(--muted);margin:.4rem 0 0}
.species-card .credit{margin:0;padding:.35rem 1.1rem .9rem}
.species-card .credit a:first-child{display:inline;color:var(--lime);text-decoration:underline}
.rows li{border-top:1px solid var(--line)}
.rows li:last-child{border-bottom:1px solid var(--line)}
.rows li>a{display:flex;justify-content:space-between;align-items:center;gap:.5rem 1rem;padding:.85rem .4rem;color:var(--text);text-decoration:none}
.rows li>a:hover h3{text-decoration:underline;text-underline-offset:.18em}
.rows p{color:var(--muted);margin:0}
.row-tags{display:flex;align-items:center;justify-content:end;flex-wrap:wrap;gap:.25rem}
.tag{display:inline-block;color:var(--lime);background:#27331e;border-radius:99rem;padding:.15rem .6rem;font-size:.79rem;font-weight:650}
.amber{color:var(--amber);background:#38291a}
.note{padding:1.1rem 1.25rem;border:1px solid var(--line);border-radius:.6rem;background:var(--surface)}
.note p:last-child{margin-bottom:0}
.note.alert{border-left:4px solid var(--amber)}
.alert .eyebrow{color:var(--amber)}
.spaced{margin-top:2.4rem}
.morph-group{padding-top:1.25rem;border-top:1px solid var(--line)}
.morph-group h3{font-size:clamp(1.4rem,3vw,1.9rem)}
.morph-group>p{max-width:72ch;color:var(--muted);margin-bottom:1rem}
.group-links,.link-list{display:flex;flex-wrap:wrap;gap:.45rem .75rem;padding:0;list-style:none}
.link-list a,.group-links a{display:inline-block;background:var(--surface);border:1px solid var(--line);border-radius:.35rem;padding:.3rem .7rem}
.profile{max-width:800px}
.profile section{padding:1.45rem 0;border-top:1px solid var(--line)}
.facts,.copies{display:grid;grid-template-columns:repeat(auto-fit,minmax(min(100%,13rem),1fr));gap:.6rem}
.facts div{padding:.55rem .75rem;background:var(--surface);border-radius:.35rem}
.facts dt,.evidence{font-size:.82rem;color:var(--muted)}
.facts dd{margin:0;font-weight:650}
.references li{padding:.55rem 0;border-bottom:1px solid var(--line)}
.references span{display:block;color:var(--muted);font-size:.82rem}
.foot{border:0;border-top:1px solid var(--line);margin-top:3.5rem;color:var(--muted);font-size:.83rem}
.foot p{margin:0}
@media(max-width:670px){.hero,.hero.split{grid-template-columns:1fr;gap:1rem}.hero-index{display:none}.bar{padding-block:.6rem}.rows li>a{align-items:start;flex-direction:column}.row-tags{justify-content:start}}